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Boraginaceae

life science Maturity 11-13 Next article →

The Boraginaceae family consists of diverse flowering plants. You may recognize them as borage or forget-me-nots. This family is part of a larger group of plants called asterids. Scientists study these plants to understand how they grow and evolve. The way we group them has changed as we learn more. Some experts use a narrow definition called Boraginaceae sensu stricto. This group includes about 90 to 120 genera. Other experts use a broader definition called Boraginaceae sensu lato. This older view includes about 155 genera.

Many plants in this family share specific physical traits. Their leaves are often arranged alternately along the stem. Some plants may have a mix of alternate and opposite leaves. The leaf blades are usually narrow, such as linear or lance shapes. The edges of the leaves can be smooth or toothed. Most species have bisexual flowers, though some taxa are dioecious. This means some plants have separate male and female individuals.

A very common feature is the presence of plant hairs. Most members of the Boraginaceae family have hairy leaves. These hairs are coarse because of cystoliths. Cystoliths are tiny structures made of silicon dioxide and calcium carbonate. These hard parts can cause skin reactions in some people. Gardeners who handle these plants regularly might experience itching or a rash. This is a direct result of the physical structure of the hairs.

The flowers of Boraginaceae follow a fascinating biological process. Many species have inflorescences that form a coiling shape when new. Scientists call this shape a scorpioid cyme. The flower itself usually has a five-lobed calyx. The corolla, or the petals, can be bell-shaped, tubular, or rotate. Colors include white, yellow, orange, pink, purple, blue, or green. Most pollination is performed by hymenopterans, such as bees.

Color changes in the flowers provide important biological signals. In some species, anthocyanins cause flowers to change from red to blue. Anthocyanins are the pigments responsible for these colors. This change happens as the flower ages. It may serve as a signal to pollinators. The color tells insects that the flower is old. It signals that the nectar and pollen are depleted.

Taxonomy, or the science of naming and grouping, has shifted significantly. The APG IV system from 2016 classified Boraginaceae as the only family in the order Boraginales. Previously, the Cronquist system placed them in the order Lamiales. However, researchers found they were not as similar to Lamiales as once thought. A 2016 revision of the Boraginales split the family into 11 distinct families. These include Codonaceae, Ehretiaceae, Hydrophyllaceae, and others. This change helped scientists group plants more accurately based on their true relationships.

There are many well-known examples of plants in this family. Alkanet and borage are very common. You might also see comfrey, fiddleneck, or forget-me-not in various environments. Other members include hound's tongue, lungwort, and the oysterplant. The purple viper's bugloss is also known as Salvation Jane. These plants show the incredible variety within a single family. From sea-dwelling oysterplants to garden comfrey, they occupy many different niches.

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