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Lamiales

life science Maturity 11-13 Next article →

The Lamiales are a massive order of flowering plants. Scientists often call them the mint order. They belong to the asterids clade within the Eudicots. This group is incredibly diverse and widespread. Under the APG IV classification system, the order contains 24 different families. It includes about 23,810 species and 1,059 genera. You can find these plants in almost every habitat on Earth. They grow in forests, valleys, and grasslands. They also live in rainforests, marshes, and even frozen areas.

Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087
Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087

Most Lamiales plants have very specific physical features. They are eudicots with either herbaceous or woody stems. A common feature is zygomorphic flowers. This term means the flowers have bilateral symmetry. These flowers usually have five petals. They often form an upper lip with two petals and a lower lip with three petals. Some species have actinomorphic flowers, which are radially symmetrical. These plants may have five stamens, but they are often reduced to two or four. They also produce a single style attached to an ovary. This ovary usually contains two carpels and is mostly superior. Many species also have glandular hairs on their surfaces.

Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087
Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087

Within this order, plants have developed very different ways to survive. Some species are carnivorous, meaning they catch and eat small animals. The families Lentibulariaceae and Byblidaceae include carnivorous plants. Other members, like those in the Martyniaceae family, are protocarnivorous. Some Lamiales are also parasitic. This means they live on or derive nutrients from other plants. The family Orobanchaceae contains these parasitic species. They can be hemi-parasites or holoparasites.

Utricularia aurea 8 Darwiniana
Utricularia aurea 8 Darwiniana
Cordylanthusrigidus
Cordylanthusrigidus

Understanding the history of Lamiales is a complex task for scientists. In the past, the order had a much more restricted circumscription. For example, Arthur Cronquist included families like Lamiaceae and Verbenaceae. The Dahlgren classification system placed them in the superorder Lamiiflorae. Recent phylogenetic work, which studies evolutionary relationships, has changed everything. Research shows the Lamiales are polyphyletic compared to the order Scrophulariales. Because of this, the two groups are now combined. This new, larger group also includes the former orders Hippuridales and Plantaginales. The name Lamiales is now the preferred term for this combined group. The placement of the Boraginaceae family remains unclear.

Utricularia aurea 8 Darwiniana
Utricularia aurea 8 Darwiniana

Scientists are still working to date the exact age of the Lamiales lineage. This is a difficult problem to solve. A study in 2004 used molecular phylogenetic dating to estimate their age. That study suggested a stem lineage of 106 million years. However, a similar study in 2009 provided a different estimate. That study suggested the lineage was 80 million years old. Another study in 2009 explained why these dates are so hard to determine. Determining the exact timing of plant evolution requires very precise data.

Cordylanthusrigidus
Cordylanthusrigidus

Humans rely on Lamiales for many essential things. Many species are economically important for food and medicine. We use aromatic herbs like basil, mint, rosemary, and sage. Other culinary plants include oregano, thyme, and marjoram. Some species provide wood, such as ash and teak. Others are used for decoration, like lilacs and jasmine. Many plants in this order produce medicinal compounds. These include alkaloids and saponins. These substances can help treat infections, asthma, or the flu. They also possess antioxidant properties.

Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087
Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087

These plants also play a role in protecting environments. Some species can repel insects. This can help control diseases like malaria, which is spread by mosquitoes. The family Acanthaceae is especially interesting in this regard. Their mature leaves contain bioactive secondary metabolites. These chemicals are toxic to insect larvae. Because of this, botanical insecticides derived from these plants are useful. They are inexpensive and abundant. They are also safer for the environment and non-target organisms than synthetic chemicals. Even the color of these plants is useful. In Sardinia, people use Lavandula stoechas to create a light-green natural dye.

Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087
Lavandula angustifolia - Köhler–s Medizinal-Pflanzen-087
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