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Anthracotheriidae

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Anthracotheriidae is a paraphyletic family of extinct animals. This means the group does not consist of a single lineage sharing one common ancestor. These animals were artiodactyl ungulates, which are even-toed hoofed mammals. They are closely related to modern hippopotamuses and whales. Understanding these animals helps scientists trace the history of how different species evolved over millions of years.

Microbunodon
Microbunodon

In life, an average anthracothere likely resembled a skinny hippopotamus. They also had a general appearance that was somewhat similar to a pig. Their heads were relatively small and narrow compared to their bodies. To move through their environments, they used broad feet with four or five toes on each foot. These wide feet were very useful for walking on soft mud without sinking. Their mouths contained full sets of about 44 teeth. Some species had upper molars with five semicrescentric cusps. These teeth were likely adapted for digging up the roots of aquatic plants.

The fossils of these animals provide clues about how they lived. The types of sediments where fossils are found suggest that anthracotheres were amphibious. This means they likely spent time both on land and in the water. This lifestyle supports the idea that they may be ancestral to modern hippopotamuses. However, the exact relationship is still a subject of scientific study. One study suggests that a different group called palaeochoerids might be the true stem group of the Hippopotamidae family.

Microbunodon
Microbunodon

Scientists have discovered anthracothere fossils in several important locations. The family gets its name from the first genus discovered, Anthracotherium. This name means "coal beast" because the first fossils were found in Paleogene-aged coal beds in France. Other significant discoveries were made in northern Pakistan. A joint research project between Harvard University and the Geological Survey of Pakistan found remains on the Pothohar Plateau. These fossils were located in well-dated middle and late Miocene deposits.

The history of the anthracotheres spans many geological periods. The oldest known genus is Elomeryx, which first appeared in Asia during the middle Eocene. Most of these animals thrived across Africa and Eurasia. During the Oligocene, a few species even moved into North America. However, they eventually disappeared from Europe and Africa during the Miocene. This disappearance may have been caused by climatic changes. They also likely faced competition from other artiodactyls, such as pigs and hippopotamuses. The youngest genus, Merycopotamus, died out in Asia during the late Pliocene, possibly for similar reasons.

Researchers use various methods to understand how anthracotheres relate to other animals. Comparative gene sequencing has shown that living hippos are the closest relatives of whales. This connection places them in a group called Whippomorpha. However, the timeline presents a puzzle for scientists. Cetaceans, which include whales and dolphins, originated about 50 million years ago in the Tethys Sea. In contrast, the Hippopotamidae family is only 15 million years old. The first Asian hippopotamids appeared only 6 million years ago. Because the anthracotheres appeared in the middle Eocene, they lived well after whales had already become fully aquatic.

There are different scientific theories regarding where anthracotheres fit in the tree of life. While many see them as stem hippos, other analyses suggest different connections. One study proposes that anthracotheres are part of a clade that includes entelodonts. This group would be a sister clade to other cetancodonts. In this theory, a genus called Siamotherium is considered the most basal member of the Cetacodontamorpha clade. These complex relationships show how much work remains to fully resolve the history of these extinct mammals.

Microbunodon
Microbunodon
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