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Artiodactyla

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Artiodactyla is a large order of placental mammals known as even-toed ungulates.

Hand skeletons with Artiodactyls highlighted
Hand skeletons with Artiodactyls highlighted

These animals are defined by how they carry their weight. They bear weight equally on an even number of toes, usually two. Specifically, they use the third and fourth toes, which often form a hard hoof. This distinguishes them from perissodactyls, which bear weight on an odd number of toes.

Walia ibex illustration white background
Walia ibex illustration white background

This group includes many animals that are vital to human culture and economy. They range from small deer to massive giraffes and camels.

One of the most important features of many artiodactyls is how they digest food. Most members of this order are herbivores that eat plants. To process tough plant cellulose, they use one or more specialized stomach chambers. This process is more efficient than the intestinal digestion used by perissodactyls. However, not all artiodactyls follow this rule. The Suina, which includes pigs, are omnivores that eat both plants and meat.

Warthog Face 001
Warthog Face 001

Cetaceans, such as whales and dolphins, are entirely carnivorous.

Modern science has revealed a surprising connection within this group.

Humpback Whale underwater shot
Humpback Whale underwater shot

Molecular biology has shown that cetaceans are part of the artiodactyl branch. Specifically, whales and dolphins are most closely related to hippopotamuses. Because of this, some taxonomists use the name Cetartiodactyla to include both groups. This ensures the group is monophyletic, meaning it includes a common ancestor and all its descendants. Other researchers use the term "even-toed ungulates" to refer only to the land-dwelling species.

The evolutionary history of artiodactyls began in the early Eocene, about 53 million years ago.

Anoplotherium commune quadrupedal
Anoplotherium commune quadrupedal

Fossils from this era appeared almost at once in Europe, Asia, and North America. This makes it difficult for scientists to pinpoint their exact place of origin. The earliest known family is Diacodexeidae. Their most famous member was Diacodexis. These were small, slim animals, sometimes only the size of a hare. They had long tails and hind legs that were longer than their front legs.

Many extinct families once roamed the Earth.

entelodon Sp Illustration
entelodon Sp Illustration

The Entelodontidae were stocky animals with massive heads and bony bumps on their lower jaws. They lived from the middle Eocene to the early Miocene in Eurasia and North America. Another group, the Anthracotheriidae, had a pig-like build and lived from the middle Eocene to the Pliocene. These animals are thought to be the ancestors of modern hippopotamuses. Hippos appeared later, in the late Miocene, and live in Africa and Asia today.

Hippopotamus amphibius
Hippopotamus amphibius

Other groups show how these animals spread across the globe. The Merycoidodontidae were a diverse group of North American camels. They lived in North America until they went extinct around 10,000 years ago.

Camelus dromedarius in Singapore Zoo
Camelus dromedarius in Singapore Zoo

Terrestrial artiodactyls only reached South America in the Pliocene. This happened after the Isthmus of Panama formed about three million years ago. Because of this late arrival, South America has fewer artiodactyl families than other continents. Australia has no native terrestrial species, and Antarctica has no terrestrial mammals at all.

Scientists confirmed these relationships using molecular biology.

Thomas Herbert Maguire - Richard Owen 1850
Thomas Herbert Maguire - Richard Owen 1850

By sequencing DNA and RNA, researchers compared genetic material between species. In 1994, scientists like Dan Graur and Desmond Higgins used these methods to link cetaceans to artiodactyls. Fossil evidence from Pakistan also provided crucial clues.

Indohyus BW
Indohyus BW

Fossils of Pakicetus, an ancient whale from 48 million years ago, showed a specific ankle bone structure. This double-rolled joint surface is a trait shared with even-toed ungulates. This physical link proves that whales evolved from land-dwelling ancestors.

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🖼️ Images & Media (29)
File:entelodon Sp Illustration.jpg
entelodon Sp Illustration.jpg
File:MEPAN Sivatherium.jpg
MEPAN Sivatherium.jpg
File:Anoplotherium commune quadrupedal.png
Anoplotherium commune quadrupedal.png
File:Thomas Herbert Maguire - Richard Owen 1850.png
Thomas Herbert Maguire - Richard Owen 1850.png
File:Humpback Whale underwater shot.jpg
Humpback Whale underwater shot.jpg
File:Hippopotamus amphibius.JPG
Hippopotamus amphibius.JPG
File:Recherches pour servir à l'histoire naturelle des mammifères (Pl. 80) (white background).jpg
Recherches pour servir à l'histoire...
File:Voyage en Abyssinie Plate 2 (white background).jpg
Voyage en Abyssinie Plate 2 (white background).jpg
File:Cladogram of Cetacea within Artiodactyla (Camelus bactrianus).png
Cladogram of Cetacea within Artiodactyla...
File:Tragulus napu - 1818-1842 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg
Tragulus napu - 1818-1842 - Print -...
File:Walia ibex illustration white background.png
Walia ibex illustration white background.png
File:Mesonyx.jpg
Mesonyx.jpg

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