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Tapejaridae

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Tapejaridae is a family of azhdarchoid pterosaurs that lived during the Cretaceous period. Their name comes from a Tupi word meaning "the lord of the ways." These flying reptiles were spread across much of the globe. Scientists have found their fossils in Africa, Asia, and Europe. They are also well-known from South America. Some evidence suggests they may have lived in North America as well.

Jiufotang tapejarids
Jiufotang tapejarids

Most tapejarids were small to medium-sized animals. They possessed several unique physical traits, especially regarding their skulls. Most species had a bony crest rising from the snout. This crest was formed mostly by the premaxillary bones at the tip of the upper jaw. In some species, this bone supported a much larger crest. This larger structure was made of soft, fibrous tissue that extended back along the skull.

Tapejarines mmartyniuk
Tapejarines mmartyniuk

The skull of a tapejarid also featured a large nasoantorbital fenestra. This is a main opening in the skull located in front of the eyes. In this family, the opening spanned at least half the length of the entire skull. Their eye sockets were small and pear-shaped. Studies of their brain cases show they had extremely good vision. This vision was better than in other pterosaur groups. They likely relied almost exclusively on sight to hunt or interact with others.

Unnamed pterosaur
Unnamed pterosaur

Their body structure was also very unusual. Tapejarids had reduced shoulder girdles that hung low on the torso. This resulted in a "bottom decker" wing arrangement. This means their wings protruded from near the belly rather than the back. This setup is reminiscent of how some modern planes are designed.

Thalassodromeus in Japan
Thalassodromeus in Japan

Scientists believe tapejarids were arboreal, meaning they lived in trees. They had more curved claws than other azhdarchoid pterosaurs. They are often found in fossil sites alongside other arboreal flying vertebrates, such as early birds. Many experts speculate they were frugivores or omnivores. This idea comes from their parrot-like beaks. Direct evidence for plant-eating exists in a Sinopterus specimen. That fossil contains phytoliths and gastroliths in its abdominal cavity.

Jiufotang tapejarids
Jiufotang tapejarids

The classification of this family has changed many times. Brazilian paleontologist Alexander Kellner named the family in 1989. He defined it as the clade containing Tapejara and Tupuxuara. In 2007, Kellner divided the family into Tapejarinae and Thalassodrominae. However, other scientists challenged this model. In 2003, David Unwin suggested that Tupuxuara and Thalassodromeus were more distantly related to Tapejara. He believed they belonged to the Azhdarchidae family instead.

Thalassodromeus in Japan
Thalassodromeus in Japan

Other researchers have proposed different groupings over the years. In 2011, a study suggested the family Chaoyangopteridae should be part of Tapejaridae. This was not followed by all authors. In 2014, Brian Andres and his colleagues defined the family as the clade containing Tapejara and Sinopterus. They also created the group Tapejarini. More recently, in 2020, a new subfamily called Sinopterinae was named. This group includes species more closely related to Sinopterus than to Tapejara.

Jiufotang tapejarids
Jiufotang tapejarids

In 2023, Rodrigo Pêgas and his team studied these relationships again. They argued that Thalassodromidae and Tapejaridae should be separate families. They found that these groups belong within a larger group called Tapejaromorpha. In 2024, Pêgas provided further updates to these names. His work helps clarify how species like Caupedactylus and Aymberedactylus fit into the tree of life. These constant changes show how much we are still learning about these ancient creatures.

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🖼️ Images & Media (4)
File:Tapejarines mmartyniuk.png
Tapejarines mmartyniuk.png
File:Jiufotang_tapejarids.png
Jiufotang_tapejarids.png
File:Thalassodromeus in Japan.jpg
Thalassodromeus in Japan.jpg
File:Unnamed pterosaur.jpg
Unnamed pterosaur.jpg
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