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Pith

life science Maturity 11-13 Next article →

Pith, also known as the medulla, is a specialized tissue found within the stems of vascular plants. It is primarily composed of soft, spongy parenchyma cells. These cells serve several biological roles, such as storing starch for the plant. The location and structure of pith vary depending on the type of plant. In eudicotyledons, the pith is situated at the very center of the stem. In monocotyledons, however, the pith extends only into the roots. The tissue is organized in a specific sequence within the plant's anatomy. The pith is encircled by a ring of xylem, which is the tissue that moves water. This xylem is then surrounded by an outer ring of phloem, which transports nutrients.

Taxus wood
Taxus wood

As plant tissue matures, its physical properties often undergo noticeable changes. New pith growth is typically white or a very pale color. As the tissue ages, it commonly darkens to a deeper brown shade. In many trees, pith is a prominent feature of young growth. However, as a tree develops an older trunk or larger branches, the pith is often replaced by xylem. This replacement process means the center of an old tree is quite different from a sapling. In some plant species, the pith in the middle of the stem may dry out. When this happens, the tissue disintegrates and leaves the stem hollow.

Walnut pith
Walnut pith

Some plants exhibit much more complex pith structures than others. For example, walnut trees possess what is known as chambered pith. This structure contains numerous short cavities within the tissue. In certain plants, the cells on the outer edges of the pith develop differently than the inner cells. This specific layer is called the perimedullary region of the pithamus. One species that shows this distinct cell development is Hedera helix, commonly known as ivy. These variations in cell structure show how specialized plant tissues can become.

Elder pith
Elder pith

Humans have found several ways to utilize pith for survival and industry. In Melanesia and Micronesia, the pith of the sago palm is a vital food source. This is due to the high starch content found within the tissue. However, the raw pith is highly toxic to animals and humans. To make it safe, people use a process to leach the toxins away from the starch. This extraction process allows only the safe starch component to be consumed. The resulting starch is very similar in appearance to tapioca. Currently, these extraction processes are generally about 50% efficient. This means half of the material remains as residual pith waste.

Elder pith
Elder pith

Beyond food, pith has served important roles in historical technology and maintenance. In the field of watchmaking, pith wood is used as a specialized cleaning tool. Watchmakers use pieces of pith, such as from the elder or mullein plants, to clean their equipment. The spongy nature of the wood allows it to remove oil from the tips of small tools. This prevents oil from contaminating the delicate movements inside a watch. Additionally, the spongy wood of the pith wood plant was once used to manufacture pith helmets. Although often mistakenly called pith, this material provided a lightweight and protective structure.

Elder pith
Elder pith

Historical lighting methods also relied on the unique properties of pith. From the 17th century until the mid-20th century, people used rushlights for home lighting. This method involved using the dried pith from certain rush plants. This pith is actually the center of the leaf. To create a flame, the dried pith was soaked in fat or grease. This setup allowed for a steady, albeit small, source of light. This method even saw a brief period of revival during World War 2.

Elder pith
Elder pith

The term "pith" is also applied to different biological structures in other contexts. In botany, it can refer to the pale, spongy inner layer of the rind in citrus fruits. This layer is more accurately called the mesocarp or the albedo. You can see this in fruits like oranges, which belong to the hesperidia group. The linguistic history of the word provides a clue to its meaning. It originates from the Old English word "piþa," which means substance. This is closely related to the Middle Dutch word "pitte," which refers to the pit of a fruit.

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File:Walnut pith.jpg
Walnut pith.jpg
File:Taxus wood.jpg
Taxus wood.jpg
File:Elder pith.jpg
Elder pith.jpg
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