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Osteoglossiformes

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Osteoglossiformes is a unique order of ray-finned fish. Their name comes from Ancient Greek words meaning "bony tongues." This name refers to their specialized, hard tongues. These fish are considered a primitive group. This means they represent an older lineage of fish. Most of the at least 245 living species live in freshwater. They are found across South America, Africa, Australia, and southern Asia.

These fish possess a very unusual internal anatomy. In almost all other fish, the digestive tract follows a specific path. For most species, the gastrointestinal tract passes to the right of the stomach. In Osteoglossiformes, this is different. The forward part of their gut passes to the left of the oesophagus and stomach. This anatomical difference is a key way to identify them. It shows how much they vary from other ray-finned fish.

Scientists divide this group into different families and suborders. The two main sub-orders are the Osteoglossoidei and the Notopteroidei. One family is the Pantodontidae, which includes freshwater butterfly fishes. Another is the Osteoglossidae, known as the bonytongues. The Arapaimidae family contains the large arapaimas. The Notopteridae family includes featherfin knifefishes or featherbacks. Within this group, there are subfamilies like the Asiatic featherbacks. There are also the African knifefishes, known as Xenomystinae.

Some members of this order have developed amazing ways to sense their world. The Mormyridae family contains elephantfishes. The Gymnarchidae family includes the African knifefish, or *Gymnarchus niloticus*. These are known as weakly electric fish. They do not use electricity to shock others. Instead, they create electric fields to sense their prey. This allows them to find food in different environments. It is a specialized way to navigate and hunt.

The history of these fish is tied to the movement of the Earth. They first evolved on the supercontinent called Gondwana. This massive landmass eventually broke apart into smaller continents. This explains why they are found in disconnected places like Africa and South America. For a long time, people thought they were only freshwater fish. However, research in 2008 changed this view. Scientists described new marine species from the Danish Eocene Fur Formation. This discovery showed that the group is even more diverse than once thought.

The size range within Osteoglossiformes is truly massive. The smallest member is *Pollimyrus castelnaui*. This tiny fish is only 1 cm long. On the other end of the scale is the arapaima, or *Arapaima gigas*. This giant fish can grow to be as long as 3 meters. This shows how much the group has changed over time. They can be tiny or enormous, depending on their specific family.

Studying these fish helps scientists understand the history of life. They provide clues about how ancient continents like Gondwana were shaped. Their unique digestive systems show how biology can take different paths. Even the mooneyes, or Hiodontidae, are often linked to this group. Some scientists believe they might belong to a separate order called Hiodontiformes. Whether in freshwater or ancient marine fossils, these fish tell a complex story of evolution.

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