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Keystone species

life science Maturity 9-11 Next article →

A keystone species is an organism that has a disproportionately large effect on its natural environment relative to its abundance. This means that even if the species does not have a huge amount of biomass or productivity, its presence is vital. The term describes how a single species can influence the types and numbers of many other organisms in a community. Without these species, an ecosystem might change dramatically or cease to exist altogether. The concept is often used in conservation biology to help protect important parts of nature.

Keystone species
Keystone species

The name comes from an analogy to a keystone in a stone arch. In an arch, the keystone is the central stone at the very top. While it may experience the least pressure of all the stones, the entire structure will collapse if it is removed. Similarly, an ecosystem can experience a massive shift if a keystone species is lost. Even if the species is just a small part of the total living matter, it holds the system together. This concept helps ecologists communicate with policy-makers about which animals need the most protection.

Zoologist Robert T. Paine introduced this concept in 1969. He developed the idea through observations and experiments in the intertidal zone. This is the area of the shoreline between the high and low tide lines. Paine studied the relationships between marine invertebrates, specifically starfish and mussels, in Makah Bay, Washington. He used the ochre starfish, or Pisaster ochraceus, as a primary example. These starfish are generalist predators that eat many things, including snails, barnacles, and limpets. However, their favorite food is the mussel, which is a dominant competitor for space on the rocks.

Keystone species
Keystone species

Paine's experiment showed exactly how the mechanism of a keystone predator works. He removed the ochre starfish from a specific area to see the results. Initially, the rock pools held 15 different rock-clinging species. After three years, only 8 species remained in the pools. After ten years, the mussels had grown so much that they occupied almost the entire space. By eating the mussels, the starfish allowed other species like seaweeds, sponges, and anemones to coexist. Without the predator, the mussels crowded everyone else out.

Keystone species
Keystone species

Many keystone species are apex predators, meaning they are at the top of the food chain. For example, the gray wolf in the Greater Yellowstone Ecosystem exerts a profound influence on the trophic pyramid. When wolves were removed, herbivores began to over-graze woody plants. This affected the entire plant population and even hurt beaver populations. Without wolves to keep them moving, herbivores grazed too heavily in riparian areas, which are the banks of rivers. When wolves were reintroduced, the riparian ecosystem and the beaver population recovered quickly.

Fig. 1 -Riparian willow recovery (26485120926) horiz
Fig. 1 -Riparian willow recovery (26485120926) horiz

Other predators also act as keystones by protecting habitats. Sea otters in the North American west coast protect kelp forests by eating sea urchins. When sea otters were hunted for fur, their numbers fell below 1,000 in the North Pacific. Without otters, urchins grazed the kelp holdfasts so heavily that the forests disappeared. In Central and South America, the jaguar helps balance the mammalian jungle ecosystem. The jaguar has a very wide diet, consuming 87 different species of prey.

Junior-Jaguar-Belize-Zoo
Junior-Jaguar-Belize-Zoo

Some species are keystones because they are ecosystem engineers. These animals physically change their environment. The beaver is a famous example of an ecosystem engineer. Beavers cut down trees to build dams, which transforms streams into ponds or wetlands. These new habitats benefit many species, including amphibians, salmon, and songbirds.

American Beaver
American Beaver

Similarly, prairie dogs in North America create tunnel systems. These burrows provide homes for owls and mountain plovers. The tunnels also help rainwater reach the water table and prevent soil erosion.

PrairieDogTownGregg
PrairieDogTownGregg

Finally, some species are keystones through mutualism, which is a mutually beneficial interaction. In Western Australia, the acorn banksia provides nectar for honeyeaters. Because the tree is often the only nectar source available, the birds depend on it. In return, the birds help pollinate many plant species. If this one tree species were lost, the honeyeater population might collapse. This would have profound implications for the entire ecosystem.

Banksia prionotes 3 gnangarra
Banksia prionotes 3 gnangarra

This shows how even a single plant can support a massive web of life.

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🖼️ Images & Media (7)
File:Junior-Jaguar-Belize-Zoo.jpg
Junior-Jaguar-Belize-Zoo.jpg
File:American Beaver.jpg
American Beaver.jpg
File:Keystone species.jpg
Keystone species.jpg
File:Fig. 1 -Riparian willow recovery (26485120926) horiz.jpg
Fig. 1 -Riparian willow recovery...
File:Banksia prionotes 3 gnangarra.jpg
Banksia prionotes 3 gnangarra.jpg
File:PrairieDogTownGregg.jpg
PrairieDogTownGregg.jpg
File:BeaverDam 8409.jpg
BeaverDam 8409.jpg
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