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Jaw

life science Maturity 11-13 Next article →

Jaws are articulated structures located at the entrance of the mouth.

Siebert 12 (jaws)
Siebert 12 (jaws)

They are used to grasp and manipulate food for many different animals. In most species, the jaws are a key part of the body plan. They serve to open and close the mouth vault. This allows animals to acquire, carry, and process food through mastication, which is the process of chewing.

Bullant head detail
Bullant head detail

In arthropods, such as insects, the jaws are made of a material called chitin. These jaws often oppose each other laterally. They may be called mandibles or chelicerae. Some of these mouthparts are actually modified legs that help move food.

Vertebrates use a different system for their jaws.

Great white shark at his back11
Great white shark at his back11

These jaws are usually made of bone or cartilage. They typically consist of an upper jaw and a lower jaw that oppose each other vertically. Most vertebrate jaws also bear numerous teeth. In some specialized fish, the jaw structure is extremely complex. Teleost fish often have jaws modified for suction feeding or jaw protrusion. This involves dozens of different bones working together. Some fish, like the moray eel, even possess two distinct sets of jaws.

Pharyngeal jaws of moray eels
Pharyngeal jaws of moray eels

The oral jaws capture the prey first. Then, the pharyngeal jaws move the prey from the mouth toward the esophagus for swallowing.

Evolutionary history suggests that jaws appeared during the Silurian period. They were first seen in Placoderm fish. These fish then diversified during the Devonian period. Scientists believe jaws evolved from the most anterior two pharyngeal arches. These are the arches that originally supported the gills. One arch became the jaw itself. The other became the hyoid arch. The hyoid system suspends the jaw from the skull's braincase. This connection allows the jaws to have great mobility. While there is no direct fossil evidence, this theory explains the number of arches seen in living animals. Jawless vertebrates, known as Agnatha, have nine arches. Jawed vertebrates, called Gnathostomes, have seven arches.

Interestingly, the first advantage of jaws might not have been eating. Jaws may have originally helped with respiration, or breathing. They could have functioned as a buccal pump. This is a mechanism that pumps water across the gills of fish. In amphibians, this same pump moves air into the lungs. Over a long period of evolutionary time, the use of jaws for feeding became the primary function. This shift provided a major selective advantage for vertebrates. The ability to chew and grab food changed how animals interacted with their environments.

Tetrapods, which include amphibians, reptiles, and birds, have simplified jaw structures. Many of the upper jaw bones are fused to the braincase. These bones include the premaxilla, maxilla, jugal, quadratojugal, and quadrate. The lower jaw bones are also fused into a single unit called the mandible. This unit includes the dentary, splenial, angular, surangular, and articular bones. The jaw moves through a hinge joint between the quadrate and the articular. Some animals have very little movement between these bones. However, snakes show a high degree of cranial kinesis. This means their skull bones move significantly, allowing them to swallow very large prey.

Mammals have a unique jaw arrangement. The mammalian jaw consists of the maxilla (the upper jaw) and the mandible (the lower jaw). During mammalian evolution, two specific bones changed significantly. The articular bone of the lower jaw and the quadrate bone were reduced in size. These two bones were actually incorporated into the ear. Many other jaw bones fused together to create a strong unit. Because of this, mammals show little to no cranial kinesis. Instead, the mandible attaches to the temporal bone via the temporomandibular joints. Dysfunction in these joints can cause pain or clicking movements.

Some animals have developed entirely different jaw systems. Sea urchins possess unique jaws that show five-part symmetry. This structure is known as Aristotle's lantern. Each unit of this lantern holds a single tooth. These teeth are made of crystalline calcium carbonate. Unlike human teeth, these teeth grow perpetually. This specialized mechanism allows sea urchins to feed in their specific marine environments. From the complex suction jaws of fish to the specialized lanterns of sea urchins, the jaw is a versatile tool of survival.

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🖼️ Images & Media (4)
File:Siebert 12 (jaws).jpg
Siebert 12 (jaws).jpg
File:Bullant head detail.jpg
Bullant head detail.jpg
File:Great white shark at his back11.jpg
Great white shark at his back11.jpg
File:Pharyngeal jaws of moray eels.svg
Pharyngeal jaws of moray eels.svg
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