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Dhole

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The dhole, known scientifically as *Cuon alpinus*, is a unique member of the canid family.

Dhole Kanha
Dhole Kanha

These animals are native to South, East, and Southeast Asia. They play a vital role in their ecosystems as specialized hunters. Currently, the dhole is listed as Endangered on the IUCN Red List. This status means their populations are steadily decreasing across their range. Experts estimate that fewer than 2,500 mature individuals remain in the wild.

Dhole in Ueno, Tokyo
Dhole in Ueno, Tokyo

Dholes are physically distinct from members of the genus *Canis*, which includes wolves and domestic dogs. Their skull profile is convex, meaning it curves outward, rather than being concave or sunken. Their dental structure is also very different. They lack a third lower molar tooth entirely. Additionally, their upper molars possess only a single cusp, or sharp point.

Dholeskull
Dholeskull

This is a reduction from the two to four cusps found in many other canids. These specialized teeth have developed into sharply trenchant, or slicing, points to aid in feeding.

Social structure is a defining feature of dhole life. They live in highly social clans rather than following a rigid dominance hierarchy.

Asiatic Wild Dogs (43518944044)
Asiatic Wild Dogs (43518944044)

While a typical clan consists of about 12 individuals, some large groups have been known to exceed 40 members. Dholes are diurnal, which means they are active and hunt during daylight hours. They preferentially target medium and large-sized ungulates, which are hoofed mammals.

2012-bandipur-dhole-sambar
2012-bandipur-dhole-sambar

In tropical forests, they must compete with large predators like the tiger (*Panthera tigris*) and the leopard (*Panthera pardus*). Although they share some prey, they often target different species to minimize this competition.

The history of the dhole involves a massive geographical range. During the Pleistocene epoch, dholes ranged throughout much of Asia.

Cuon alpinus Cova Negra
Cuon alpinus Cova Negra

Their range even extended into Europe, and one controversial record suggests they may have reached North America. However, their range became much more restricted about 12,000 to 18,000 years ago. Today, they are extinct in Central Asia and parts of Southeast Asia. They may also be extinct on the Korean peninsula and in parts of Russia.

Scientific study of the dhole has evolved significantly over the centuries. In 1811, Peter Simon Pallas proposed the binomial name *Canis alpinus*. He described a range that reached much further north into Russia than where they live today. In 1833, Brian Houghton Hodgson suggested the name *Canis primaevus*. He believed the dhole was a primitive form and a progenitor, or ancestor, of the domestic dog.

Later, Hodgson recognized their physical differences and proposed the genus *Cuon*. In 1955, paleontologist Erich Thenius concluded they descended from a golden jackal-like ancestor.

Recent genetic research has provided new insights into dhole evolution. In 2018, whole genome sequencing revealed evidence of ancient genetic admixture between the dhole and the African wild dog (*Lycaon pictus*).

This suggests the two species may have interbred in the past, possibly in the Middle East. While they are closely related to the *Canis* genus, they also share an ancient ancestry with the *Lycaon* lineage. This relationship is even seen in the extinct Sardinian dhole (*Cynotherium sardous*), which is most closely related to the living dhole among all canines.

There are several recognized subspecies of dholes, though genetic studies show no clear distinctions between them.

Keulemans Cuon alpinus javanicus
Keulemans Cuon alpinus javanicus

One major grouping includes populations from South and Central India into Myanmar. Another group extends from northern India through Thailand and the Malaysian Peninsula. The origins of dholes in Sumatra and Java remain unclear.

Cuon alpinus alpinus puppy
Cuon alpinus alpinus puppy

Some researchers speculate these island populations may have been introduced by humans. The survival of the species depends on addressing threats like habitat loss, loss of prey, and disease transfer from domestic dogs.

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🖼️ Images & Media (29)
File:Cuon alpinus Cova Negra.jpg
Cuon alpinus Cova Negra.jpg
File:Leopold v. Schrenck - Cuon alpinus.png
Leopold v. Schrenck - Cuon alpinus.png
File:202104 Dog.svg
202104 Dog.svg
File:Dogs, jackals, wolves, and foxes (Plate I).jpg
Dogs, jackals, wolves, and foxes (Plate I).jpg
File:Dogs, jackals, wolves, and foxes (Plate IX).jpg
Dogs, jackals, wolves, and foxes (Plate IX).jpg
File:Dogs, jackals, wolves, and foxes (Plate XI).jpg
Dogs, jackals, wolves, and foxes (Plate XI).jpg
File:Dogs, jackals, wolves, and foxes (Plate X).jpg
Dogs, jackals, wolves, and foxes (Plate X).jpg
File:Dogs, jackals, wolves, and foxes (Plate VI).jpg
Dogs, jackals, wolves, and foxes (Plate VI).jpg
File:Dogs, jackals, wolves, and foxes (Plate XLI).jpg
Dogs, jackals, wolves, and foxes (Plate XLI).jpg
File:Dogs, jackals, wolves, and foxes (Plate XLIV).jpg
Dogs, jackals, wolves, and foxes (Plate XLIV).jpg
File:Dogs, jackals, wolves, and foxes (Plate XIII).jpg
Dogs, jackals, wolves, and foxes (Plate XIII).jpg
File:Dogs, jackals, wolves, and foxes (Plate XII).jpg
Dogs, jackals, wolves, and foxes (Plate XII).jpg

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