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Arthropod

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Arthropods are invertebrates belonging to the phylum Arthropoda. This group is the largest animal phylum on Earth. Scientists estimate there are between 1.17 million and 10 million different species. This means arthropods account for over 80 percent of all known living animals.

They are vital to our world because they act as pollinators for many crops. However, some species can also spread diseases to humans and livestock. They are among the few animal groups to successfully adapt to dry land.

The defining feature of an arthropod is its exoskeleton. This is a hard outer shell made of a polymer called chitin. The cuticle, or outer layer, usually consists of three distinct parts. First is the epicuticle, a thin waxy coat that prevents moisture loss. Below that is the exocuticle, which contains chitin and hardened proteins. The innermost layer is the endocuticle, made of chitin and unhardened proteins.

Arthropod cuticle based upon Xvazquez edited to include english legend
Arthropod cuticle based upon Xvazquez edited to include english legend

In many crustaceans and beetles, the shell is biomineralized with calcium carbonate. This makes the armor even tougher and stronger for defense.

Because an exoskeleton is rigid, it cannot stretch as the animal grows. To increase in size, arthropods must undergo ecdysis, also known as moulting. During this process, the animal sheds its old shell, called an exuviae. A new, softer exoskeleton is revealed underneath. The animal then grows until it is time to moult again.

Cicada climbing out of its exoskeleton while attached to tree
Cicada climbing out of its exoskeleton while attached to tree

This cycle continues until the arthropod reaches full adult size.

Arthropod bodies are built from repeated segments. These segments can group together into specialized regions called tagmata. For example, many insects have a body divided into three parts. Spiders have a different two-part body structure.

20211214 arthropod biramous arthropodized appendage morphology en
20211214 arthropod biramous arthropodized appendage morphology en

Most arthropods have paired, jointed appendages. These limbs are highly specialized, acting like a Swiss Army knife. Some appendages function as gills for breathing, while others serve as claws, antennae, or walking legs. In some species, these limbs have even vanished entirely.

Sensory systems in arthropods are incredibly diverse. Most use a combination of compound eyes and pigment-pit ocelli. While compound eyes provide most visual information, ocelli often detect light direction. Spiders are a notable exception. Their main eyes are ocelli that can actually form images. Some spider eyes can even swivel to track moving prey.

Arthropoda montage (eyes)
Arthropoda montage (eyes)

They also use setae, which are tiny bristles projecting through the cuticle. These setae act as mechanical or chemical sensors to detect touch or currents.

Internal systems also show great variety. Most arthropods have an open circulatory system. Instead of veins, they have a body cavity called a haemocoel. A fluid called haemolymph circulates through this cavity to reach internal organs. Their nervous systems are ladder-like. They feature paired ventral nerve cords that run through every segment. These cords form clusters of nerve cells called ganglia in each segment. In the head, these ganglia fuse to form the brain, which encircles the esophagus.

The history of arthropods stretches back to the Cambrian period. Scientists study fossils like Marrella to understand their early ancestors.

Marrella
Marrella

The name "arthropod" comes from the Greek words for "jointed leg." The term was first used in anatomical descriptions by Barthélemy Charles Joseph Dumortier in 1832. In 1843, the German zoologist Johann Ludwig Christian Gravenhorst first used the designation "Arthropoda." Determining exactly how all these different body types evolved is known as "The arthropod head problem."

There is an enormous scale of size within this phylum. The smallest arthropods are crustaceans in the class Tantulocarida, some measuring less than 1 mm. In contrast, the largest are found in the class Malacostraca. The Japanese spider crab has legs that can span up to 3.7 meters. The American lobster can reach weights exceeding 20 kg.

Whether they are microscopic or massive, arthropods remain a dominant force in every ecosystem on Earth.

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🖼️ Images & Media (60)
File:Protaetic cuprea ignicollis 2023-03-22 IZE-066 zoom.jpg
Protaetic cuprea ignicollis 2023-03-22...
File:20211214 arthropod biramous arthropodized appendage morphology en.png
20211214 arthropod biramous arthropodized...
File:Arthropod cuticle based upon Xvazquez edited to include english legend.PNG
Arthropod cuticle based upon Xvazquez...
File:Cicada climbing out of its exoskeleton while attached to tree.jpg
Cicada climbing out of its exoskeleton...
File:Pone.0028183.g005.png
Pone.0028183.g005.png
File:20200213 Remipedia Nectiopoda central nervous system.png
20200213 Remipedia Nectiopoda central...
File:Tliltocatl albopilosus 1.jpg
Tliltocatl albopilosus 1.jpg
File:Arthropoda montage (eyes).png
Arthropoda montage (eyes).png
File:Wasp ocelli.JPG
Wasp ocelli.JPG
File:Shrimp nauplius.jpg
Shrimp nauplius.jpg
File:Marrella.png
Marrella.png
File:20210310 Kylinxia zhangi.png
20210310 Kylinxia zhangi.png

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