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Scyphozoa

life science Maturity 11-13 Next article →

Scyphozoa are a class of marine animals known as the true jellyfish. They belong to the phylum Cnidaria and live exclusively in saltwater oceans. These organisms are famous for their distinct cup-like shape. This shape is reflected in their name, which comes from the Greek word *skyphos*, meaning a drinking cup.

Stranded Cambrian scyphozoans
Stranded Cambrian scyphozoans

Scyphozoans have a very long history on Earth. Fossil evidence shows they have existed since the earliest Cambrian period. While they are often seen in coastal waters, they live throughout the world's oceans. They can be found anywhere from the surface to great depths. However, they never live in freshwater or on land.

The biology of a Scyphozoan involves a complex life cycle with two main phases. The first phase is a bottom-dwelling polyp. This stage is often inconspicuous and can live for a long time. The second phase is the planktonic medusa. The medusa is the bell-shaped form most people recognize.

Schleiden-meduse-2
Schleiden-meduse-2

The life cycle begins when a fertilized egg produces a planular larva. This larva quickly attaches to the sea floor and becomes a tiny, stationary polyp called a scyphistoma. The scyphistoma can reproduce asexually by budding new polyps. Eventually, the polyp undergoes strobilation. This is a process where the polyp buds off many tiny medusae from its upper surface. These young medusae start out microscopic and may take years to reach sexual maturity.

Anatomy in Scyphozoans is quite different from many other animals. They have no durable hard parts. This means they have no head, no skeleton, and no specialized organs for breathing or excretion. Instead, they rely on mesoglea for structural integrity. Mesoglea is an internal gelatinous material that acts like a skeleton. This material contains mobile amoeboid cells that come from the epidermis. Most Scyphozoans also display four-part symmetry. To move, they use a ring of muscle fibers located within the mesoglea around the rim of the dome. They swim by alternately contracting and relaxing these muscles. This rhythmic movement allows them to catch prey or escape from predators.

Feeding in Scyphozoans relies on specialized stinging cells called nematocysts. These cells are located on the tentacles that hang from the edge of the umbrella dome. They are also found on the four or eight oral arms that hang from the central mouth. Most Scyphozoans use these cells to capture crustaceans and fish. Some species are instead filter feeders. They use their tentacles to strain tiny plankton from the water. The mouth opens into a central stomach. From this stomach, four interconnected diverticula radiate outward. In many species, a system of radial canals helps distribute nutrients. Some genera, such as *Cassiopea*, even have smaller mouths located on their oral arms.

Even without a brain, Scyphozoans can sense their environment. Their nervous system is usually a distributed net of cells. Some species have more organized nerve rings. In species without these rings, nerve cells are concentrated into small structures called rhopalia. There are between four and sixteen of these lobes arranged around the rim of the umbrella. The rhopalia coordinate the muscular actions needed for swimming. Each rhopalium is typically connected to a pair of sensory pits. They also contain a statocyst and sometimes a pigment-cup ocellus. These parts help the animal stay oriented and detect light.

Scyphozoans vary greatly in size and impact. Most species range from a few centimeters to a few meters in diameter. The species *Cyanea capillata* can reach very large widths. Some jellyfish are commercially important, such as the moon jelly (*Aurelia aurita*). Others, like the enormous *Nemopilema nomurai*, can cause major disruptions to fisheries. This large species is found between Japan and China. Because jellyfish can consist of as much as 98% water, they are rarely preserved as fossils. However, we know they are diverse. While scientists recognize about 200 extant species, the true diversity is likely at least 400 species.

The classification of Scyphozoa has changed over time. They were once thought to include the classes Cubozoa and Staurozoa. Now, they include only three extant orders. Two of these orders are part of the subclass Discomedusae. The other major group is the subclass Coronamedusae. This complex system of life shows how simple structures can perform many vital tasks. From the sea floor polyp to the drifting medusa, Scyphozoans connect different parts of the ocean ecosystem.

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File:Stranded Cambrian scyphozoans.jpg
Stranded Cambrian scyphozoans.jpg
File:Schleiden-meduse-2.jpg
Schleiden-meduse-2.jpg
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