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Procyonidae

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Procyonidae is a family of mammals found in the New World. This group belongs to the order Carnivora. It includes many well-known animals like raccoons, coatis, and ringtails. It also includes kinkajous, olingos, and olinguitos.

While they are part of the carnivore order, most procyonids are omnivores. This means they eat a wide variety of both plants and animals. Because they eat many different things, they have adapted to live in many different environments. This flexibility makes them a very successful group of animals.

Most procyonids have specific physical traits that define them. They generally have slender bodies and long tails. The common raccoon is an exception because it tends to be more bulky. Many people view them as small cousins of the bear family. This is reflected in German names like Waschbär, which means washing bear. Other names include Nasenbär for coatis and Honigbär for kinkajous.

ZooKeys - Bassaricyon medius (white background)
ZooKeys - Bassaricyon medius (white background)

However, science shows they are actually more closely related to mustelids. They share traits like a shortened rostrum, which is the snout area. They also lack alisphenoid canals and have a relatively flat mandibular fossa.

Their teeth are specialized for their omnivorous diets. Most species have a dental formula that totals 40 teeth. The kinkajou is unique because it has only 36 teeth. This is because it has one fewer premolar in each row. Kinkajous also have special features for living in trees. They have a prehensile tail, which means the tail can grasp objects. They also have a unique femoral structure in their legs.

ZooKeys - Bassaricyon alleni (white background)
ZooKeys - Bassaricyon alleni (white background)

Because they eat many things, they have lost some meat-eating adaptations. Their carnassial teeth, which are used for shearing meat, are poorly developed in most species. This is especially true for the common raccoon.

Procyonids have a very deep evolutionary history. Fossil evidence shows ancestors lived during the Miocene epoch. These fossils date back approximately 20 million years (Ma). Some fossil groups, like Stromeriella, once lived in the Old World. However, they went extinct during the Pliocene.

ZooKeys - Bassaricyon gabbii (white background)
ZooKeys - Bassaricyon gabbii (white background)

It is suggested that early procyonids were an offshoot of canids. They likely adapted to an omnivorous diet over time. They also moved into South America during the Great American Interchange. This event began about 7.3 Ma ago in the late Miocene. The genus Cyonasua was part of this movement.

Genetic studies have helped scientists map their family tree. Kinkajous are a sister group to all other living procyonids. They split from the others about 22.6 Ma ago. The branch leading to coatis and olingos separated from the raccoon branch about 17.7 Ma ago. Later, olingos and coatis diverged about 10.2 Ma ago. This happened around the same time ringtails and raccoons parted ways. The separation between coatis and mountain coatis occurred about 7.7 Ma ago.

ZooKeys - Bassaricyon neblina (white background)
ZooKeys - Bassaricyon neblina (white background)

These dates help us understand how the family grew.

Social structures vary greatly across the family. Most members of Procyonidae are solitary animals. However, some species prefer to live in groups. Female coatis form bands of 4 to 24 individuals. These bands forage together to find food. Kinkajous also show social behavior in small groups. These groups might consist of two males and one female. Reproduction patterns also differ. Ringtails, olingos, and kinkajous typically give birth to one offspring. Raccoons and coatis have litters ranging from 2 to 6 offspring.

Classification of these animals has changed as technology improved. Scientists once thought the red panda was a procyonid. Molecular biology studies now place it in the family Ailuridae.

ZooKeys - Bassaricyon alleni (white background)
ZooKeys - Bassaricyon alleni (white background)

In the past, olingos were often grouped together as one subspecies. DNA sequence data proved they were actually different. Older maps grouped kinkajous and olingos together based on appearance. We now know this was parallel evolution. This means they developed similar traits independently. Modern genetic analysis provides a much more accurate view of their true relationships.

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File:Wild animals of North America, intimate studies of big and little creatures of the mammal kingdom (Page 410) (white background).jpg
Wild animals of North America, intimate...
File:ZooKeys - Bassaricyon medius (white background).jpg
ZooKeys - Bassaricyon medius (white...
File:ZooKeys - Bassaricyon alleni (white background).jpg
ZooKeys - Bassaricyon alleni (white...
File:ZooKeys - Bassaricyon gabbii (white background).jpg
ZooKeys - Bassaricyon gabbii (white...
File:ZooKeys - Bassaricyon neblina (white background).jpg
ZooKeys - Bassaricyon neblina (white...
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