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Pholcidae

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The Pholcidae are a large family of araneomorph spiders found on every continent except Antarctica. This family includes more than 2,000 individual species. They are organized into 94 different genera, which are groups of related species. People often call them by many names, such as cellar spiders, daddy long-legs, or vibrating spiders.

Cellar Spider preying upon house spider
Cellar Spider preying upon house spider

While they share the name "daddy long-legs" with harvestmen and crane flies, they are biologically very different. Harvestmen have only one pair of eyes and a single body segment. In contrast, pholcids are true spiders with two distinct body segments.

Pholcids have a very specific physical structure. Their bodies consist of a prosoma, the front section, and an opisthosoma, the rear section. The prosoma might be shaped like a dome or have a small pit in the center. The opisthosoma varies greatly in shape; it can be long and cylindrical, short, or even pointed at the back. Most pholcids have eight eyes arranged in two groups of three, sometimes with a small middle pair. However, some genera like Modisimus and Spermophora only have six eyes. Their legs are famously long and thin, featuring flexible tarsi, which are the segments at the very end of the legs.

These spiders are masters of a unique hunting method. Unlike many other spiders, pholcid webs have no adhesive or sticky properties. Instead, the web is built in a messy, irregular shape to trap prey. When a pholcid detects an insect in its web, it quickly envelops the prey in silk-like material. The spider may eat the meal immediately or store it for later. Once the spider is finished feeding, it cleans the web by unhooking the remains and letting the carcass drop away.

Two Crossopriza lyoni egg bundle magnified
Two Crossopriza lyoni egg bundle magnified

When a pholcid feels threatened, it uses a specialized behavior called a threat response. If something touches the web, the spider begins to vibrate or gyrate rapidly. This movement can sometimes settle into a circular rhythm. The spider might oscillate in tune with the elasticity of the web, creating a motion larger than its own legs. This behavior has earned them the name "vibrating spiders." Scientists believe this might confuse predators or signal to rivals. It may also help the spider capture insects that have just brushed against the web.

Pholcids have a very interesting diet that includes other spiders. While they primarily eat insects, certain species will invade the webs of other spiders. They may eat the host spider, its eggs, or its captured prey. Some species even mimic the vibrations of struggling prey to lure a host spider closer. They are known to prey on Tegenaria funnel weaver spiders, redback spiders, huntsman spiders, and house spiders. Because they eat certain species, they may actually be beneficial to humans by keeping spider populations in check.

The history of this family was first established by Carl Ludwig Koch in 1850. Since then, researchers have identified many different species across the globe. Their size varies significantly depending on the species. For example, Spermophorides lascars is just over 1 millimeter long. Meanwhile, Artema atlanta can reach a body length of 11 mm. They are highly adaptable and can be found in dark, damp places like caves, under rocks, or in abandoned mammal burrows. In human homes, they prefer undisturbed areas like attics and cellars.

There is a common urban legend regarding the danger of these spiders. Many people believe pholcids possess the most potent venom of any spider, but that their fangs are too weak to bite humans. This is a misconception. Pholcids do have a short, hooked fang structure called an uncate. While they can bite humans, their venom is not medically significant and is relatively weak. In 2004, the television show MythBusters tested this. They found the venom was much less toxic than that of a black widow. One host even experienced a bite that caused only a mild, short-lived burning sensation.

The legend likely exists because pholcids successfully hunt much more venomous spiders, such as the redback or members of the Latrodectus genus.

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🖼️ Images & Media (4)
File:Two Crossopriza lyoni egg bundle magnified.jpg
Two Crossopriza lyoni egg bundle magnified.jpg
Crossopriza lyoni Video.ogv
File:Spermophora senoculata pedipalps.jpg
Spermophora senoculata pedipalps.jpg
File:Cellar Spider preying upon house spider.jpg
Cellar Spider preying upon house spider.jpg
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