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Phloem

life science Maturity 9-11 Next article →

Phloem is a specialized living tissue found in vascular plants. Its primary role is to transport organic compounds produced during photosynthesis. These compounds are known as photosynthates, and the most common is the sugar sucrose. This movement of nutrients is called translocation. In many plants, the phloem is located as the innermost layer of the bark. This is why the name comes from the Ancient Greek word "phloiós," meaning bark.

Xylem and phloem diagram
Xylem and phloem diagram

Translocation works through a process of phloem loading and unloading. The movement is driven by positive hydrostatic pressures. This differs from the xylem, which uses negative pressures or tension. The sap is a water-based solution rich in sugars. It moves from a "source" to a "sink." A source is a part of the plant that produces food, such as a leaf. A sink is a part that consumes or stores food, such as a root, a bulb, or a developing fruit.

Translocation from the source to the sink within the phloem
Translocation from the source to the sink within the phloem

During the spring growth period, storage organs like roots often act as sources. These provide energy to the many growing areas of the plant, which act as sinks. Once the growth period ends and meristems become dormant, the leaves become the main sources. In this stage, the storage organs and developing seeds become the sinks. Because the flow is multidirectional, sap can move up or down. This means sap in adjacent sieve tubes might even flow in opposite directions.

Phloem and Xylem in stem
Phloem and Xylem in stem

Phloem tissue is composed of several distinct cell types working together. The main conducting cells are called sieve tube elements. To allow fluids to move easily, these cells lack a nucleus and most organelles at maturity. They do contain vacuoles and ribosomes before they mature, but these migrate to the cell wall and dissolve. The ends of these cells have pores called sieve areas. These pores are reinforced by platelets of a polysaccharide called callose.

Phloem cells
Phloem cells

Because sieve tube elements lack a nucleus, they depend on companion cells. These are specialized parenchyma cells that handle the metabolic needs of the sieve tubes. There are three types of companion cells. Ordinary companion cells have smooth walls. Transfer cells have highly folded walls to help scavenge solutes using energy. Intermediary cells possess many vacuoles and help synthesize certain oligosaccharides. In some plants, like gymnosperms, albuminous cells perform a similar role to companion cells.

Stem-histology-cross-section-tag
Stem-histology-cross-section-tag

Other cells in the phloem provide mechanical support through sclerenchyma. This includes long, narrow fibres and irregularly shaped sclereids. Fibres provide tensile strength and flexibility. Sclereids add compression strength and can protect against herbivores. For example, sclereids create the gritty texture found in pears. The term "phloem" was introduced by Carl Nägeli in 1858. He used it to distinguish the tissue formed outwardly by the cambium from the xylem.

Stem-histology-cross-section-tag
Stem-histology-cross-section-tag

Understanding phloem is vital for studying plant survival and agriculture. If the bark is removed in a ring, it is called girdling. This process kills the plant because nutrients cannot reach the roots. However, farmers use girdling to produce massive fruits. By removing all but one fruit on a branch, all the branch's sugar is directed to that single piece. Historically, people in Finland and Sweden used pine phloem during famines. They dried and milled it into a flour called pettu to make dark bread.

Detaching inner bark of pine
Detaching inner bark of pine
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🖼️ Images & Media (6)
File:Xylem and phloem diagram.svg
Xylem and phloem diagram.svg
File:Stem-histology-cross-section-tag.svg
Stem-histology-cross-section-tag.svg
File:Phloem cells.svg
Phloem cells.svg
File:Phloem and Xylem in stem.svg
Phloem and Xylem in stem.svg
File:Translocation from the source to the sink within the phloem.svg
Translocation from the source to the sink...
File:Detaching inner bark of pine.jpg
Detaching inner bark of pine.jpg
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