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Ossicles

life science Maturity 11-13 Next article →

The auditory ossicles are three irregular bones located within the middle ear of humans and other mammals. The term ossicle comes from the Latin word for "tiny bone." These structures are among the smallest bones found in the human body. They function as a kinematic chain, which is a series of connected parts that move together. Their primary purpose is to transmit and amplify sound vibrations. These vibrations are collected from the air by the eardrum. The ossicles then move these vibrations into the fluid-filled labyrinth, or cochlea, of the inner ear. If these bones are missing or damaged, a person may experience conductive hearing loss.

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The ossicles consist of the malleus, the incus, and the stapes. They are arranged in a specific order from the eardrum to the inner ear. The malleus, which means "hammer" in Latin, is the first bone in the chain. It is attached to the tympanic membrane, also known as the eardrum. When sound hits the eardrum, the malleus moves. The malleus connects to the incus, or "anvil," through the incudomalleolar joint. The incus is the middle bone and connects to both the malleus and the stapes. The stapes, which means "stirrup," is the final bone in the sequence. It is the smallest bone in the entire human body. The stapes connects to the incus through the incudostapedial joint. It then attaches to the membrane of the fenestra ovalis, which is the oval window. This window is the opening between the middle ear and the vestibule of the inner ear.

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The way these bones move is a vital process called impedance matching. Sound traveling through air mostly reflects when it hits a liquid medium. In fact, only about 1/30 of the sound energy moving through the air would transfer into a liquid. This is why sound seems to stop abruptly when a person's head is submerged underwater. The ossicles solve this problem by providing a mechanical advantage through lever action. They also reduce the area over which the force is distributed. This process makes the vibrations stronger, even though they move a shorter distance. By increasing the pressure, the ossicles allow most of the sound energy to enter the liquid of the cochlea. This efficient coupling ensures the stimulus is successfully transmitted to the inner ear.

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Two specific muscles are attached to the ossicles to control their movement. These are the tensor tympani and the stapedius muscles. These muscles can contract to dampen the vibrations of the bones. Scientists believe this happens for two possible reasons. One theory is that this protects the inner ear from excessively loud noises. A second theory suggests that these muscles help with frequency resolution. They may do this by reducing the transmission of low frequencies, which helps the ear process higher frequencies better. These muscles are much more highly developed in bats. In bats, these muscles help block out the animal's own loud cries used during echolocation.

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The history of discovering these bones involves many different anatomists from the 16th century. There is some doubt about who discovered them first. Some researchers attribute the discovery to Alessandro Achillini or Jacopo Berengario da Carpi. In 1521, Berengario da Carpi provided the first written description of the malleus and incus. He noted they might be associated with transmitting sound. In 1536, Niccolo Massa described the bones in more detail and called them little hammers. Andreas Vesalius provided a very detailed description in 1543. He was the first person to compare the incus to an anvil. The stapes was described later in 1549 by Pedro Jimeno. However, Giovanni Filippo Ingrassia had described it in public lectures in 1546.

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Evolutionary studies show that these bones have a fascinating origin. In mammal embryos, the ear bones are actually attached to the dentary, which is part of the lower jaw. They begin as ossified portions of cartilage known as Meckel's cartilage. As the embryo develops, this cartilage hardens into bone. Later in development, the bone structure breaks away from the jaw. The bones then migrate to the middle ear area. This structure in mammals corresponds to different parts of the jaw in birds, reptiles, or amphibians. In those animals, the equivalent structures are known as the columella, quadrate, articular, and angular bones.

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Understanding the ossicles is also important for modern medicine. Sometimes the joints between these bones can become rigid. One specific condition is called otosclerosis. In this condition, the stapes fuses to the oval window. This fusion reduces a person's ability to hear. This medical issue can sometimes be treated with surgery. Doctors may use a passive middle ear implant to help restore hearing. This shows how the precise movement of these tiny bones is essential for the complex system of human hearing.

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