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Galaxiidae

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The Galaxiidae are a family of mostly small freshwater fish. They are primarily found in the Southern Hemisphere. Most species inhabit Southern Australia or New Zealand. However, some live in South Africa and South America. You can also find them on Lord Howe Island, New Caledonia, and the Falkland Islands. One specific species, the common galaxias (Galaxias maculatus), is very important. It is likely the most widely distributed natural freshwater fish in the Southern Hemisphere.

Many galaxiids follow a complex life cycle known as amphidromy. This process involves moving between fresh water and the ocean. First, the larvae hatch within a river or stream. These larvae are then washed downstream into the sea. Once in the ocean, they exist as part of the zooplankton. Later, they return to rivers as juveniles to finish their development. They eventually grow into full adults in the fresh water.

Scientists categorize the family into seven different genera. There are approximately 50 species within the Galaxiidae family. The largest group is the genus Galaxias, which contains 34 species. Other genera include Aplochiton, which has two species, and Lovettia, which has only one. The genus Neochanna includes six species, while Galaxiella and Paragalaxias each have four. The genus Brachygalaxias contains two species. There is also the disputed genus Stompooria from South Africa.

Evolutionary history suggests these fish have very ancient origins. Their ancestors likely diverged from neoteleosts around the Triassic-Jurassic boundary. While their origins are old, the earliest definitive fossils are from the Miocene of New Zealand. These fossils belong to the existing genus Galaxias. In 1998, Anderson described a possible Late Cretaceous fossil named Stompooria. Some researchers question this because Stompooria has scales. Others suggest it might be a sister group to salmoniforms.

Geography plays a major role in how these fish evolved. Many genera show unusual relationships between distant continents. For example, the Australian genus Lovettia is closely related to the South American Aplochiton. Scientists believe an Antarctic land bridge influenced this pattern. This bridge provided a shallow marine and fresh water connection. It allowed different taxa to disperse between Australia and South America.

Today, many galaxiid species face serious threats from exotic species. Humans have introduced salmonids, such as trout, to many different land masses. These trout prey upon galaxiids and compete with them for food. This has led to many localized extinctions in various places. In Australia, some species like the barred galaxias are considered threatened. In New Zealand, trout have changed the dominance of native fish.

Humans also interact with galaxiids through fishing. In some regions, the juveniles are caught as whitebait. These young fish are considered a valued delicacy. In New Zealand, exploitation of adults may be banned. This protection often applies unless the fish is for indigenous tribes. In Australia, some anglers ignore galaxiids because they are small. However, some species are large enough to be caught with fly-fishing tackle.

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