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Fly

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Flies are highly diverse insects belonging to the order Diptera. The name Diptera refers to their unique flight mechanism. These insects use only a single pair of wings for aerial movement.

Common house fly, Musca domestica
Common house fly, Musca domestica

Their hindwings have evolved into specialized mechanosensory organs called halteres. These club-like structures sense rotation during flight. This sensory system allows flies to perform advanced and precise aerobatics. As a major insect order, Diptera includes more than 150,000 described species. This group includes familiar insects like mosquitoes, houseflies, and crane flies.

The anatomy of a fly is specialized for survival and movement. A fly's body consists of three main sections called tagmata. The first is the head, which contains the sensory organs. It features large compound eyes and often three small simple eyes called ocelli. The ocelli detect changes in light intensity to help the fly react to objects.

Anthomyiidae (female) (10144905255)
Anthomyiidae (female) (10144905255)

The second section is the thorax, which houses the flight muscles. The third section is the abdomen, which consists of 11 segments. The final three segments are often modified for reproduction.

Flies possess specialized mouthparts designed for specific feeding habits. Some groups, such as mosquitoes and robber flies, have mouthparts for piercing and sucking. Female horse-flies use knife-like mandibles and maxillae to make incisions in skin. They then lap up the resulting blood. Other flies have mouthparts meant for lapping and sucking liquids.

Musca domestica female
Musca domestica female

Their guts include large structures called diverticulae. These allow the insect to store liquid after a meal. To help them navigate, flies use motion-sensitive neurons to analyze visual information. These neurons allow them to estimate self-motion and separate objects from the background.

Most flies undergo a process called complete metamorphosis, or holometabolism. This involves radical changes through several distinct life stages. First, flies lay eggs, often on a specific food source. These eggs hatch into larvae, which are commonly called maggots.

Lutzomyia adiketis
Lutzomyia adiketis

These larvae lack true limbs and often develop in protected environments. Some species are ovoviviparous, meaning they deposit hatched larvae instead of eggs. After the larval stage, the insect becomes a pupa. The pupa is a tough capsule that protects the developing insect. Finally, the adult fly emerges from this capsule.

The evolutionary history of Diptera spans hundreds of millions of years. The first true dipterans appeared during the Middle Triassic, around 240 million years ago.

Fossil insect Diptera, Brachycera in Baltic amber. Age 50 Mill. years (the Lower Eocene)
Fossil insect Diptera, Brachycera in Baltic amber. Age 50 Mill. years (the Lower Eocene)

Because flowering plants did not exist until the Cretaceous, early flies likely ate different foods. Scientists suggest they may have fed on honeydew from sap-sucking bugs. The fossil record shows three major episodes of evolutionary radiation. The first occurred in the Triassic about 220 million years ago. A second radiation of lower Brachycera happened in the Jurassic, around 180 million years ago. A third radiation of Schizophora began 66 million years ago.

Flies are ecologically significant as both helpers and pests. They are the second largest group of pollinators, following only the Hymenoptera.

In wet or cold environments, they are even more important than bees for pollination. However, some flies cause significant harm to humans and animals. Mosquitoes act as vectors for diseases like malaria, dengue, and yellow fever. Houseflies can spread foodborne illnesses globally. Larger flies, such as tsetse flies, cause economic damage to cattle. Conversely, some larvae are used in medicine for debridement to clean wounds.

Diversity within the order is immense and varies by region. There are approximately 19,000 species in Europe and 23,000 in the Oriental region.

The size of flies varies wildly across the order. The Gauromydas heros is considered the largest fly in the world. In contrast, the Euryplatea nanaknihali is smaller than a grain of salt. Flies are found in nearly all terrestrial habitats. Some species even live in extreme environments like glaciers, hot springs, or even crude oil. This wide range of adaptations makes them one of the most successful insect groups on Earth.

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🖼️ Images & Media (42)
File:Anthomyiidae (female) (10144905255).jpg
Anthomyiidae (female) (10144905255).jpg
File:European wasp white bg.jpg
European wasp white bg.jpg
File:Corydalus cornutus illustration (rotated).png
Corydalus cornutus illustration (rotated).png
File:Osmylus_(white_background).jpg
Osmylus_(white_background).jpg
File:Pseudacrossus przewalskyi (Reitter, 1887).jpg
Pseudacrossus przewalskyi (Reitter, 1887).jpg
File:Elenchus koebelei.jpg
Elenchus koebelei.jpg
File:RHYACOPHILA DORSALIS Male Pont Forge de Sailly Watigny 02 MHNT.jpg
RHYACOPHILA DORSALIS Male Pont Forge de...
File:Arctia villica SLU.JPG
Arctia villica SLU.JPG
File:Common house fly, Musca domestica.jpg
Common house fly, Musca domestica.jpg
File:Scorpionfly (white background).jpg
Scorpionfly (white background).jpg
File:Pulex irritans female ZSM (white background).jpg
Pulex irritans female ZSM (white background).jpg
File:Fossil insect Diptera, Brachycera in Baltic amber. Age 50 Mill. years (the Lower Eocene).jpg
Fossil insect Diptera, Brachycera in...

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