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Finger

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A finger is a prominent digit located on the forelimbs of most tetrapod vertebrates. In many species, especially primates, these digits are prehensile, meaning they are capable of grasping. Most tetrapods follow a pattern known as pentadactyly, which means they possess five digits.

Scheme human hand bones-en
Scheme human hand bones-en

While shorter digits on the hindlimbs are typically called toes, the elongated digits on the forelimbs are referred to as fingers. In humans, fingers are essential for tactile sensation and the fine motor movements required for dexterity.

The internal structure of a human finger is composed of bones called phalanges. Each hand contains fourteen phalanges in total. The thumb is unique because it contains only two phalanges: a proximal and a distal phalanx. The other four fingers each contain three phalanges: the proximal, middle, and distal phalanx. These bones are connected to the metacarpal bones in the palm.

Scheme human hand bones-en
Scheme human hand bones-en

Joints form wherever these bones meet, allowing for movement. There are three main joints in each finger: the metacarpophalangeal joint at the base, the proximal interphalangeal joint in the middle, and the distal interphalangeal joint near the tip. Small ossified nodes called sesamoid bones may also be embedded in the tendons to provide leverage.

Fingers do not contain muscles themselves, with the exception of the arrector pili. Instead, movement is controlled by muscles located in the palm and the forearm. These muscles connect to the finger bones via long tendons. Muscles are categorized as either extrinsic or intrinsic. Extrinsic muscles have their muscle belly in the forearm. The long flexors allow the fingers to bend, while the extensors, located on the back of the forearm, allow the digits to straighten.

Intrinsic muscles, such as the thenar and hypothenar groups, are located within the hand itself. The thenar muscles specifically allow the thumb to move in opposition, which is vital for grasping.

Movement in the fingers is a complex coordination of flexion, extension, abduction, and adduction. Flexion, or bending, is the strongest movement. While fingers appear to move independently, they actually experience finger interdependence, also known as finger enslaving. This means moving one finger often causes slight movements in the others. The precision of these movements is clearly visible in skilled tasks, such as a pianist playing a musical piece.

This high level of control is managed by the direct corticomotoneuronal pathway in the brain.

The fingertips are among the most sensitive parts of the human body. They contain the highest concentration of touch receptors and thermoreceptors in the skin. This allows humans to sense pressure, vibration, texture, moisture, and temperature. A 2013 study even suggested that fingers can detect nano-scale wrinkles on smooth surfaces. Interestingly, fingertips wrinkle when immersed in water. This is not caused by swelling, but by the sympathetic nervous system signaling blood vessels to constrict. Some scientists propose the "rain tread" hypothesis, suggesting these wrinkles help grip wet objects.

Human fingers can also undergo remarkable biological processes, such as regeneration. In children, if a fingertip is torn off, it can regrow in less than eight weeks. This happens because the distal phalanges are regenerative in youth, and stem cells in the nails create new tissue. However, this regrowth only occurs if the injury is above the nail. There are also various clinical anomalies. Polydactyly is a condition where an individual is born with more than five digits, affecting about 1 in 500 humans.

Syndactyly type1 hands
Syndactyly type1 hands

Another condition, syndactyly, involves webbed or fused fingers. This can even affect the brain, as the somatosensory cortex maps for fused fingers appear fused as well.

Fingers serve as a vital connection between the biological body and the physical environment. They act as sensory probes that allow us to ascertain the properties of the world. Beyond simple touch, the ratio of finger lengths, such as the index and ring fingers, has been linked to different biological and behavioral traits. From the evolutionary perspective of terrestrial vertebrates, the finger structure is a highly modified version of the pectoral fins found in lobe-finned fish. This evolutionary history explains why the basic pentadactyl plan is so widely shared across different species.

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File:Scheme human hand bones-en.svg
Scheme human hand bones-en.svg
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File:Syndactyly type1 hands.jpg
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