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Ammonoidea

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Ammonoids were a diverse group of extinct cephalopods. They belonged to the subclass Ammonoidea. While they look similar to modern nautiluses, they are actually more closely related to living octopuses, squid, and cuttlefish. These living relatives belong to a group called Coleoidea. Most people refer to their fossils as ammonites. This name is most often used for the order Ammonitida.

Haeckel Ammonitida
Haeckel Ammonitida

The shell of an ammonoid is a complex structure. The chambered part of the shell is called the phragmocone. This part contains several chambers called camerae. Thin walls known as septa divide these chambers from one another. The living animal occupied only the last and largest chamber, called the body chamber. As the animal grew, it added newer, larger chambers to the open end of the coil.

Ammonite shell morphology
Ammonite shell morphology

To move through the ocean, ammonoids used a mechanism for buoyancy. A thin living tube called the siphuncle passed through the septa. This tube connected the chambers of the phragmocone to the living body. Through a process called hyperosmotic active transport, the animal emptied water out of the empty chambers. This allowed the ammonoid to control its position in the water column. By adjusting the gas and water in its shell, it could rise or sink.

Neochetoceras cf. praecursor conch and internal anatomy reconstruction
Neochetoceras cf. praecursor conch and internal anatomy reconstruction

Scientists distinguish ammonoids from nautiloids by looking at their internal anatomy. In most ammonoids, the siphuncle runs along the outer rim of the shell, known as the venter. In nautiloids, the siphuncle runs through the center of the chambers. Additionally, the septa of ammonoids have unique shapes. They often have bulges and indentations. The intersection where the septum meets the outer shell is called the suture line. These lines form patterns of peaks called saddles and valleys called lobes.

Damesites cf. damesi septa (2)
Damesites cf. damesi septa (2)

Ammonoids showed incredible variety in their physical forms. Most shells were planispirals, meaning they coiled in a flat plane. Some were involute, where the outer whorl covers the previous ones. Others were evolute, where the whorls do not cover each other. During the Cretaceous period, many heteromorphs appeared. These were ammonoids with non-spiral or helical shapes.

Heteromorph shell archaetypes
Heteromorph shell archaetypes

The history of ammonoids spans hundreds of millions of years. They first appeared during the Emsian stage of the Early Devonian, about 410 to 408 million years ago. They likely evolved from straight-shelled nautiloids called Bactritida. The group survived several mass extinctions, such as the end-Permian event. During the Jurassic and Cretaceous, they became very diverse. The order Ammonitida was a major group during this time.

Ammonoidea evolutionary history
Ammonoidea evolutionary history

Ammonoids varied greatly in size. The smallest known type was Maximites from the Upper Carboniferous. Some species were massive. The Titanites of the Jurassic reached 0.5 meters in diameter. The Parapuzosia seppenradensis of the Cretaceous was even larger. This species could reach a diameter of nearly 1 meter.

Parapuzosia seppenradensis 3
Parapuzosia seppenradensis 3

All ammonoids eventually went extinct. This occurred during the Cretaceous–Paleogene extinction event about 66 million years ago. This event was caused by the Chicxulub impact. Some scientists suggest that ocean acidification caused by the impact killed them. The larvae of ammonoids were likely small and planktonic. This would have made them very vulnerable to changing ocean chemistry. While nautiluses survived by laying eggs on the seafloor, the ammonoids vanished.

Ammonites main sutural patterns
Ammonites main sutural patterns
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🖼️ Images & Media (8)
File:Haeckel Ammonitida.jpg
Haeckel Ammonitida.jpg
File:Ammonoidea evolutionary history.PNG
Ammonoidea evolutionary history.PNG
File:Parapuzosia seppenradensis 3.jpg
Parapuzosia seppenradensis 3.jpg
File:Damesites cf. damesi septa (2).png
Damesites cf. damesi septa (2).png
File:Ammonites main sutural patterns.PNG
Ammonites main sutural patterns.PNG
File:Ammonite shell morphology.png
Ammonite shell morphology.png
File:Heteromorph shell archaetypes.png
Heteromorph shell archaetypes.png
File:Neochetoceras cf. praecursor conch and internal anatomy reconstruction.png
Neochetoceras cf. praecursor conch and...
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